The United States has just experienced the second-warmest winter on record. Viewed over a longer time horizon, however, this moment is part of a much larger story. Within a single lifetime, winter temperatures across the country have risen dramatically, illustrating the accelerating pace of environmental change. As the planet’s ecological systems shift, the institutions historically responsible for conservation are also coming under strain. These converging trends raise a critical question: what kinds of regional institutions and planning approaches will be needed to design and steward sustainable landscapes in the decades ahead?